* For all those who don't know about basic wolf pack hierarchy, I'll be explaining it here. I'll be listing the ranks in order.
First the alphas. There are only two alphas. The alphas are the absolute leaders of the pack, they make the major decisions and are often the only pair allowed to have pups. The pack will often start off with one alpha (male or female) until choosing a mate. The mate they choose becomes the other alpha. When hunting/feeding, they get the first share of meat, and whatever choice parts they wish.
Now, the betas. They are second in command to the alphas. They are often bigger and stronger than the alphas, for their job is to carry out the will of the alphas and enforce their decisions. There are only up to two betas, a male and a female, they are mates. In larger packs, the alphas may permit the betas to have pups. When hunting/feeding, they get the second share of meat, after the alphas are finished. They can choose whichever parts they want that the alphas haven't claimed.
Next come the gammas. They are third in command and help the betas enforce things, yet they are not as dominant or powerful. There are 1 or sometimes 2 gamma pairs, who are usually mates. They are VERY rarely allowed to have pups. When hunting/feeding, they eat after the betas, choosing whatever parts they want that the betas haven't taken.
After gammas, come mid-ranking wolves, sometimes called deltas. Mid-ranking wolves make up the majority of the pack, and their job is to make it look as if the pack has more members than it actually does. This discourages predators or other packs from attacking. It is also their job to protect the important members of the pack when a stressful situation arrives. There is no distinct number of mid-ranking wolves, it differs with each pack. When hunting/feeding, they eat after the gammas. This is often messy, because there are quite a few mid-ranking wolves, and they sometimes fight over choice parts.
Then, the low-ranking wolves. Usually young or old and weak wolves, they very often help take care of any pups when the pack is out hunting. They sometimes get picked on by more powerful woves, but this is not very common. There is no definate number of low-ranking wolves in a pack. When hunting/feeding, they eat after the mid-ranking wolves. There is not much choice when it gets down to the low-ranking wolves, so fighting is uncommon during feeding with this rank.
After them, comes the omega. While his position is the lowest in the pack, he is also a valuable specialist member. His job is to break up fights and lower the stress in tense situations, like feeding time. He also sometimes is left behind during hunting to watch the pups. When hunting/feeding, the omega eats last. However, in return for his services, the alphas often have saved some quality meat for the omega wolf. There is only one omega wolf, male or female.
Finally, the pups** Pups are not an actual rank, but they are below the omega wolf. The pups are left behind by the den site when the pack hunts, with either the omega, or often a low-ranking female wolf. Food is always brought back for the pups when the pack goes hunting. This meat is diveded between the pups and the wolf who was watching them.
Hunters and nannies are NOT actual ranks. They are specialized jobs/positions. Nannies are often low-ranking female wolves, or an aunt to the alphas' pups. Their job is to watch the pups while the rest of the pack hunts. Hunters consist of all of the wolves except the nanny wolf and the pups. Sometimes not all of the wolves hunt at once. The best hunters are smaller built and very swift in movement.
Note: I have done much research, and this information is taken from several reliable sources, mostly books and articles, some videos and documentaries.
**: See next post!